Wildlife of Borneo

Great Mormon

Papilio memnon

A large swallowtail whose females can look remarkably unlike one another. Field records confirm it in Sarawak and Sabah, while genetic research explains its striking female wing-pattern variation.

Species details & taxonomy
Species
Papilio memnon
Genus
Papilio
Family
Papilionidae
Order
Lepidoptera
Class
Insecta
Great Mormon — Papilio memnon
Great Mormon (Papilio memnon).Hsu Hong Lin from 南投縣集集鎮, 中華民國 · Wikimedia Commons · CC BY 2.0 · Original

Recognition

Male and female appearance differs, and females occur in mimetic and non-mimetic forms. In studied populations, mimetic females combine particular hindwing colours, tails and abdominal colour to resemble unpalatable swallowtails. This is Batesian mimicry: an edible butterfly gains protection by resembling a less palatable one. Pattern alone should be assessed alongside locality and regional identification references.

Sources & references · 1Frontiers in Insect Science · doublesex controls hindwing and abdominal mimicry traits in Papilio memnon (2022)

Habitat and Borneo range

A Bungo Range survey lists Papilio memnon memnon in Sarawak and includes a photograph. A Sabah Forestry survey also recorded it at Mokotog near Keningau. These independent records establish occurrence on Borneo. They should not be confused with captive butterfly-house records or with changing classifications of the similarly named lowii taxon.

Sources & references · 2Universiti Malaysia Sarawak · Butterflies of Bungo Range: checklist and Papilio memnon photographSabah Forestry Department technical report · Kepelbagaian Serangga di Kg. Mokotog, Sook, Keningau, Sabah (2015)

Reproduction

Breeding experiments show that alternative female forms are inherited through a region involving the doublesex gene. The female pattern is not simply a response to the flower she visits. Controlled crosses also followed caterpillars, pupae and emerging adults; growth and final body size differed between sexes. The experiments explain development, while local Bornean breeding seasons remain unestablished.

Sources & references · 2Frontiers in Insect Science · doublesex controls hindwing and abdominal mimicry traits in Papilio memnon (2022)Journal of Insect Physiology · Development and adult traits among Papilio memnon mimicry genotypes (2018)

Food and ecological role

Caterpillars have been reared on several Citrus species in Indonesian research. A food plant supplies leaves throughout the growing larval stage, whereas the flying adult uses flowers. The caterpillar also possesses an eversible scent gland behind the head, the osmeterium; chemical studies of this species show that its secretion changes during larval development.

Sources & references · 2Universitas Pendidikan Indonesia research repository · Development of Papilio memnon on three Citrus species (2015)Journal of Chemical Ecology · Larval osmeterial secretions of the swallowtails, Honda (1981)

Where to look

A recorded occurrence does not guarantee a sighting or visitor access. Read the area and experience guides for habitat, responsible observation, guiding and current access before choosing a trip. Keep wildlife at a distance and follow local restrictions.

Places with documented occurrence

Sources & references

Universiti Malaysia Sarawak · Butterflies of Bungo Range: checklist and Papilio memnon photographAccessed: 2026-10-04

Universitas Pendidikan Indonesia research repository · Development of Papilio memnon on three Citrus species (2015)Accessed: 2026-10-04

Journal of Insect Physiology · Development and adult traits among Papilio memnon mimicry genotypes (2018)Accessed: 2026-10-04

Frontiers in Insect Science · doublesex controls hindwing and abdominal mimicry traits in Papilio memnon (2022)Accessed: 2026-10-04

Journal of Chemical Ecology · Larval osmeterial secretions of the swallowtails, Honda (1981)Accessed: 2026-10-04

Sabah Forestry Department technical report · Kepelbagaian Serangga di Kg. Mokotog, Sook, Keningau, Sabah (2015)Accessed: 2026-10-04

Report a correction