Wildlife of Borneo

Bornean leaffish

Nandus nebulosus

A small predator disguised by a dead-leaf outline, the Bornean leaffish lives in streams and flooded forest. Its quiet appearance conceals a diet of small fish and crustaceans.

Least concern
Species details & taxonomy
Species
Nandus nebulosus
Genus
Nandus
Family
Nandidae
Order
Anabantiformes
Class
Actinopterygii
Size
Maximum reported total length: 12 cm
Bornean leaffish (Nandus nebulosus).
Bornean leaffish (Nandus nebulosus).R.workoran · Wikimedia Commons · CC BY-SA 4.0 · Original

Recognition

Mottled colour and an irregular leaf-like profile break up the fish’s outline. It resembles a piece of dead vegetation rather than a brightly marked open-water fish. The reported maximum total length is about twelve centimetres, with smaller individuals common in reference accounts. The leaf resemblance is its most useful first impression, while a clear view of the whole fish is better than a partly concealed head.

Sources & references · 1Species account or primary research · FishBase: Nandus nebulosus, citing Kottelat and Widjanarti (2005)

Habitat and Borneo range

Small streams, flooded forest and peat-associated waters provide its habitat. The broader range runs from Thailand to Indonesia, so the common name does not mean that it occurs only on Borneo. In Central Kalimantan, Sebangau surveys place it within a landscape of blackwater river channels and seasonal forest pools. Fallen vegetation and flooding make water and forest inseparable parts of this setting.

Sources & references · 2Species account or primary research · FishBase: Nandus nebulosus, citing Kottelat and Widjanarti (2005)Mires and Peat · Rasbora kalochroma: documented occurrence

Food and ecological role

The species is usually solitary and feeds on small fishes and crustaceans. Those animal prey distinguish its feeding role from algae-grazing gouramis in the same broader wetland fauna. Its cryptic appearance also makes it easy to overlook beside leaf litter. A quiet fish among dead leaves can therefore be an active member of the aquatic food web even when no chase is visible.

Sources & references · 1Species account or primary research · FishBase: Nandus nebulosus, citing Kottelat and Widjanarti (2005)

Where to look

A recorded occurrence does not guarantee a sighting or visitor access. Read the area and experience guides for habitat, responsible observation, guiding and current access before choosing a trip. Keep wildlife at a distance and follow local restrictions.

Places with documented occurrence

Sources & references

Mires and Peat · Rasbora kalochroma: documented occurrenceAccessed: 2026-10-01

FishBase · Nandus nebulosus: reference profileAccessed: 2026-10-01

Species account or primary research · FishBase: Nandus nebulosus, citing Kottelat and Widjanarti (2005)Accessed: 2026-10-04

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