The Pinnacles on Mount Api, Gunung Mulu National Park; above-ground karst.
The Pinnacles on Mount Api, Gunung Mulu National Park; above-ground karst.Nur Nafis Naim · Wikimedia Commons · CC BY-SA 4.0 · Cropped · Original
Limestone karst · Borneo

Limestone and karst ecosystems

Karst connects rock, water and living communities. Sarawak’s Mulu landscape shows how limestone ridges, pinnacles and underground drainage belong to one landscape while supporting different ecological settings.

Characteristics

Water dissolving limestone shapes connected surface and underground features. At Mulu, the limestone of Gunung Api and its caves provide a documented example. A pinnacles photograph illustrates an above-ground karst formation; it should not be presented as a view of a cave interior or evidence of a particular animal population.

Sources & references · 2UNESCO World Heritage Centre · Gunung Mulu National ParkBritish Geological Survey (2025) · How bat excrement is sculpting Borneo’s hidden caves

Conservation

Conservation must protect cave systems and their surface surroundings together. At Mulu, biological inputs from the forest influence cave conditions; at Niah, visitor care also protects archaeological deposits. Follow designated routes and leave rock formations intact.

Sources & references · 2British Geological Survey (2025) · How bat excrement is sculpting Borneo’s hidden cavesUNESCO World Heritage Centre · The Archaeological Heritage of Niah National Park’s Caves Complex

Species found here

Examples in the linked species profiles include Edible-nest swiftlet. Their profiles identify the particular locality and evidence. A habitat association does not mean the species occurs in every patch of that habitat.

Sources & references · 5IOC World Bird List · IOC species updates: Germain’s Swiftlet lumped with Edible-nest SwiftletJournal of Avian Biology / Price, Johnson and Clayton; University of Utah repository · The evolution of echolocation in swiftletsRaffles Bulletin of Zoology / Goh et al. / National University of Singapore · Genetic diversity among white-nest swiftlets of the genus Aerodramus of house-farms in MalaysiaNotornis 68(1) / Birds New Zealand · Arthropods in the diets of swiftlets in SabahOryx / study authors · Protection of the white-nest swiftlet in the Andaman Islands, India: an assessment

Where this habitat occurs

The Mulu and Niah records connect this habitat with two researched Sarawak landscapes. Their heritage values differ: Mulu’s recognition centres on nature, while Niah’s focuses on archaeological evidence in a limestone massif. Neither designation means every limestone exposure in Sarawak has the same species, access rules or protection boundary.

Sources & references · 2UNESCO World Heritage Centre · Gunung Mulu National ParkUNESCO World Heritage Centre · The Archaeological Heritage of Niah National Park’s Caves Complex

Protected areas

Documented landscapes to explore include Gunung Mulu National Park, Niah National Park. These areas contain habitat mosaics; the habitat does not cover their entire surface. Check each area’s access information before a visit.

Sources & references · 3UNESCO World Heritage Centre · Gunung Mulu National ParkBritish Geological Survey (2025) · How bat excrement is sculpting Borneo’s hidden cavesUNESCO World Heritage Centre · The Archaeological Heritage of Niah National Park’s Caves Complex

Ecological importance

Surface forests, streams and caves exchange water and biological inputs. Changes above ground can affect the underground environment. Karst therefore asks for attention to the whole drainage system, not only spectacular rocks. In caves, darkness and the supply of food create conditions very different from those experienced in the surface forest.

Sources & references · 2UNESCO World Heritage Centre · Gunung Mulu National ParkBritish Geological Survey (2025) · How bat excrement is sculpting Borneo’s hidden caves

Sources & references

British Geological Survey (2025) · How bat excrement is sculpting Borneo’s hidden cavesAccessed: 2026-10-02

Notornis 68(1) / Birds New Zealand · Arthropods in the diets of swiftlets in SabahAccessed: 2026-10-02

Oryx / study authors · Protection of the white-nest swiftlet in the Andaman Islands, India: an assessmentAccessed: 2026-10-02

IOC World Bird List · IOC species updates: Germain’s Swiftlet lumped with Edible-nest SwiftletAccessed: 2026-10-02

Raffles Bulletin of Zoology / Goh et al. / National University of Singapore · Genetic diversity among white-nest swiftlets of the genus Aerodramus of house-farms in MalaysiaAccessed: 2026-10-02

Journal of Avian Biology / Price, Johnson and Clayton; University of Utah repository · The evolution of echolocation in swiftletsAccessed: 2026-10-02

UNESCO World Heritage Centre · Gunung Mulu National ParkAccessed: 2026-10-02

UNESCO World Heritage Centre · The Archaeological Heritage of Niah National Park’s Caves ComplexAccessed: 2026-10-02

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